BILLY TWELVETREES has been released by the England squad ahead of the international with South Africa tomorrow and returns to skipper Gloucester as Harlequins come to Kingsholm in the Aviva Premiership tonight (7.45pm).

With James Hook having been called up by Wales, Twelvetrees will switch from his normal position as centre to play at fly half, partnering Dan Robson at half back.

Following the LV= Cup weekends, the Gloucester starting line-up has a more familiar look to it as a number of regulars return after a couple of weeks off.

In addition to Hook, there are another six players away with international teams – Jonny May and Ben Morgan (England), Greig Laidlaw (Scotland) and Sione Kalamafoni, Aleki Lutui, and Sila Puafisi (Tonga).

However, Richard Hibbard, not required by Wales this weekend, returns to his club and will start among the replacements.

Gloucester: Rob Cook; Charlie Sharples, Billy Meakes, Mark Atkinson, Henry Purdy; Billy Twelvetrees (capt), Dan Robson; Nick Wood, Darren Dawidiuk, John Afoa; Tom Palmer, James Hudson; Tom Savage, Matt Kvesic, Ross Moriarty Replacements: Richard Hibbard, James Gibbons, Yann Thomas, Elliott Stooke, Gareth Evans, Callum Braley, Aled Thomas, Dan Thomas.